:root {
  color-scheme: light;
  --bg: #ffffff; /* page background */
  --sidebar: #3f735d; /* matte green primary accent / navigation */
  --sidebar-muted: #d8e9e0; /* light text on green navigation */
  --surface: #ffffff; /* main panels and tables */
  --surface-soft: #edf6f1; /* light green secondary accent */
  --text: #1f2d27; /* primary body text */
  --muted: #61786d; /* muted helper text */
  --border: #d4e3db; /* subtle green-gray borders */
  --accent: #3f735d; /* matte green primary accent */
  --accent-light: #edf6f1; /* light green secondary accent */
  --accent-dark: #2f5a47; /* darker green hover state */
  --link: #2f5a47; /* links */
  --danger: #9f1239; /* error text */
}

* {
  box-sizing: border-box;
}

body {
  min-height: 100vh;
  margin: 0;
  background: var(--bg);
  color: var(--text);
  font-family: Inter, ui-sans-serif, system-ui, -apple-system, BlinkMacSystemFont, "Segoe UI", sans-serif;
  /* font-family: sans-serif; */
  font-size: 15px;
  line-height: 1.5;
}

a {
  color: var(--link);
}

.sidebar {
  position: fixed;
  inset: 0 auto 0 0;
  width: 228px;
  padding: 35px 18px;
  background: var(--sidebar);
  color: #ffffff;
}

.brand {
  display: flex;
  align-items: center;
  justify-content: flex-end;
  gap: 10px;
  padding-right: 10px;
  color: #ffffff;
  font-weight: 700;
  font-size: 19px;
  text-decoration: none;
}

.nav {
  display: grid;
  gap: 0;
  margin-top: 70px;
}

.nav a {
  padding: 15px 0;
  border-bottom: 1px solid rgba(255, 255, 255, 0.22);
  color: var(--sidebar-muted);
  font-weight: 600;
  text-decoration: none;
  text-align: right;
  padding-right: 10px;
}

.nav a:last-child {
  border-bottom: none;
}

.nav a:hover,
.nav a.active {
  color: #ffffff;
}

.main {
  min-height: 100vh;
  margin-left: 228px;
  padding: 36px;
}

.content {
  width: min(1180px, 100%);
}

.section {
  margin-bottom: 28px;
}

.home-section {
  width: min(820px, 100%);
}

.home-intro {
  margin-bottom: 46px;
}

.page-header {
  display: flex;
  flex-wrap: wrap;
  align-items: flex-end;
  justify-content: space-between;
  gap: 14px;
  width: min(1180px, 100%);
  margin-bottom: 18px;
  letter-spacing: 1px;
  border-bottom: 1px solid var(--border);
  padding-bottom: 8px;
}

.page-header h2 {
  margin-bottom: 0;
}

.page-header h1 {
  min-width: 0;
  overflow-wrap: anywhere;
}

.section-header {
  display: flex;
  flex-wrap: wrap;
  align-items: flex-end;
  justify-content: space-between;
  gap: 14px;
  width: min(1180px, 100%);
  margin-bottom: 18px;
  letter-spacing: 1px;
  border-bottom: 1px solid var(--border);
  padding-bottom: 8px;
}

.section-header h2 {
  margin-bottom: 0;
}

h1,
h2,
h3,
p {
  margin-top: 0;
}

h1 {
  color: var(--accent);
  font-weight: 400;
  margin-bottom: 10px;
  /* font-size: clamp(30px, 4vw, 44px); */
  font-size: 35px;
  line-height: 1.12;
}

h2 {
  color: var(--accent);
  font-weight: 400;
  margin-bottom: 12px;
  font-size: 24px;
  line-height: 1.2;
}

h3 {
  color: var(--accent);
  font-weight: 400;
  margin-bottom: 8px;
  font-size: 17px;
}

.lede {
  max-width: 820px;
  color: var(--text);
  font-size: 16px;
}

.reference-note {
  margin-bottom: 0;
}

.home-panel {
  width: min(820px, 100%);
}

.panel {
  overflow: hidden;
  border: 1px solid var(--border);
  border-radius: 2px;
  background: var(--surface);
}

.panel-body {
  padding: 18px;
}

.table-wrap {
  width: 100%;
  overflow-x: auto;
}

table {
  width: 100%;
  border-collapse: collapse;
}

th,
td {
  padding: 12px 14px;
  border-bottom: 1px solid var(--border);
  text-align: left;
  vertical-align: top;
}

th {
  background: var(--accent-light);
  color: var(--text);
  font-size: 12px;
  font-weight: 800;
  text-transform: uppercase;
}

tr:last-child th,
tr:last-child td {
  border-bottom: 0;
}

.summary-panel th {
  width: 230px;
}

.functional-summary-header {
  display: flex;
  flex-wrap: wrap;
  align-items: center;
  gap: 8px;
  width: min(1180px, 100%);
  margin-bottom: 18px;
  letter-spacing: 1px;
  border-bottom: 1px solid var(--border);
  padding-bottom: 8px;
}

.functional-summary-header h2 {
  margin-bottom: 0;
}

.info-tooltip {
  position: relative;
  display: inline-flex;
  align-items: center;
  justify-content: center;
  color: var(--muted);
  outline: none;
}

.info-icon {
  display: inline-flex;
  align-items: center;
  justify-content: center;
  width: 18px;
  height: 18px;
  border: 1px solid var(--border);
  border-radius: 50%;
  background: var(--surface);
  color: var(--muted);
  font-size: 12px;
  font-weight: 800;
  line-height: 1;
}

.tooltip-content {
  position: absolute;
  z-index: 20;
  bottom: calc(100% + 8px);
  left: 50%;
  display: none;
  width: min(420px, calc(100vw - 72px));
  padding: 10px 12px;
  border: 1px solid var(--border);
  border-radius: 2px;
  background: var(--surface);
  color: var(--text);
  box-shadow: 0 12px 26px rgba(31, 41, 51, 0.14);
  font-size: 13px;
  font-weight: 500;
  line-height: 1.45;
  text-transform: none;
  transform: translateX(-50%);
}

.tooltip-content ul {
  margin: 0;
  padding-left: 18px;
}

.tooltip-content li + li {
  margin-top: 6px;
}

.info-tooltip:hover .tooltip-content,
.info-tooltip:focus .tooltip-content {
  display: block;
}

.functional-summary-stack {
  display: grid;
  gap: 18px;
}

.functional-summary-subsection {
  display: grid;
  gap: 10px;
}

.functional-summary-subsection h3 {
  width: min(1180px, 100%);
  margin-bottom: 0;
  padding-bottom: 8px;
  /* border-bottom: 1px solid var(--border); */
  color: var(--accent);
  font-size: 19px;
  font-weight: 400;
  line-height: 1.2;
}

.functional-summary-panel {
  overflow: visible;
}

.functional-summary-panel .table-wrap {
  overflow-x: auto;
}

.functional-summary-table {
  min-width: 720px;
}

.functional-summary-table th,
.functional-summary-table td {
  vertical-align: top;
}

.functional-summary-table th {
  overflow-wrap: anywhere;
}

.functional-summary-table td.numeric {
  text-align: right;
}

.coverage-summary-table th {
  width: 340px;
  background: var(--surface);
  color: var(--text);
  font-size: 13px;
  font-weight: 700;
  text-transform: none;
  letter-spacing: 0;
}

.coverage-summary-table td {
  width: 220px;
  text-align: right;
  font-weight: 400;
}

.functional-summary-type {
  color: var(--muted);
  font-size: 13px;
  font-weight: 700;
  white-space: nowrap;
}

.functional-summary-cell {
  display: grid;
  gap: 3px;
  min-width: 0;
}

.functional-summary-primary {
  color: var(--text);
  font-weight: 400;
  line-height: 1.35;
  overflow-wrap: anywhere;
}

.functional-summary-primary .table-link,
.annotation-value .table-link,
.annotation-inline-values .table-link {
  color: var(--accent);
}

.functional-summary-id,
.annotation-id {
  white-space: nowrap;
}

.functional-summary-separator {
  color: var(--muted);
  font-size: 13px;
}

.annotation-separator {
  color: var(--muted);
  font-size: 12px;
}

.functional-summary-link-list {
  display: flex;
  flex-wrap: wrap;
  justify-content: flex-start;
  gap: 4px 10px;
}

.functional-summary-link-list .table-link {
  white-space: nowrap;
}

.functional-summary-secondary,
.functional-summary-muted {
  color: var(--muted);
  font-size: 13px;
  line-height: 1.35;
}

.functional-summary-empty {
  padding: 18px;
  border: 1px solid var(--border);
  border-radius: 2px;
  background: var(--surface);
  color: var(--muted);
  font-weight: 600;
}

.numeric {
  font-variant-numeric: tabular-nums;
  white-space: nowrap;
}

.annotation-label {
  color: var(--accent);
  font-size: 11px;
  font-weight: 800;
  letter-spacing: 0;
  text-transform: uppercase;
}

.gene-table {
  min-width: 940px;
  table-layout: fixed;
}

.gene-table th:nth-child(1),
.gene-table td:nth-child(1) {
  width: 13%;
}

.gene-table th:nth-child(2),
.gene-table td:nth-child(2) {
  width: 43%;
}

.gene-table th:nth-child(3),
.gene-table td:nth-child(3) {
  width: 15%;
}

.gene-table th:nth-child(4),
.gene-table td:nth-child(4) {
  width: 19%;
}

.gene-table th:nth-child(5),
.gene-table td:nth-child(5) {
  width: 10%;
}

.gene-table tbody td {
  border-bottom-style: dotted;
}

.gene-name span {
  display: block;
  overflow: hidden;
  text-overflow: ellipsis;
  white-space: nowrap;
}

.gene-position {
  white-space: nowrap;
}

.annotation-cell {
  white-space: normal;
}

.annotation-stack {
  display: grid;
  gap: 10px;
  max-width: 100%;
}

.annotation-section {
  display: grid;
  gap: 3px;
}

.annotation-values {
  display: grid;
  gap: 5px;
}

.annotation-entry {
  min-width: 0;
}

.annotation-value {
  overflow: hidden;
  color: var(--text);
  font-size: 13px;
  line-height: 1.35;
  text-overflow: ellipsis;
}

.annotation-value.primary {
  color: var(--text);
  font-size: 14px;
}

.annotation-secondary,
.annotation-empty,
.annotation-more {
  color: var(--muted);
  font-size: 12px;
  line-height: 1.35;
}

.annotation-secondary,
.annotation-value {
  display: -webkit-box;
  -webkit-box-orient: vertical;
  -webkit-line-clamp: 2;
  overflow: hidden;
}

.annotation-inline-values {
  display: flex;
  flex-wrap: wrap;
  gap: 4px 10px;
  color: var(--text);
  font-size: 13px;
}

.annotation-inline-values span {
  white-space: nowrap;
}

.annotation-more {
  width: max-content;
}

.annotation-more summary {
  width: max-content;
  color: var(--muted);
  font-size: 12px;
  font-weight: 700;
  cursor: pointer;
}

.annotation-more-body {
  display: grid;
  max-height: 220px;
  overflow-y: auto;
  gap: 5px;
  padding-top: 6px;
}

.annotation-more-body.inline {
  display: flex;
  flex-wrap: wrap;
  gap: 4px 10px;
}

.muted {
  color: var(--muted);
}

.button-row {
  display: flex;
  flex-wrap: wrap;
  align-items: center;
  gap: 10px;
  min-width: 0;
}

.button,
button {
  display: inline-flex;
  align-items: center;
  justify-content: center;
  min-height: 36px;
  padding: 8px 12px;
  border: 1px solid var(--border);
  border-radius: 2px;
  background: var(--surface);
  color: var(--accent);
  font: inherit;
  font-weight: 700;
  text-decoration: none;
  cursor: pointer;
}

.button.primary {
  border-color: var(--accent);
  background: var(--accent);
  color: #ffffff;
}

.button:hover,
button:hover {
  border-color: var(--accent);
}

.button.primary:hover {
  background: var(--accent-dark);
}

.button.disabled,
button:disabled,
.pager-button.disabled {
  opacity: 0.45;
  pointer-events: none;
}

.icon-button {
  display: inline-flex;
  align-items: center;
  justify-content: center;
  width: 34px;
  min-width: 34px;
  min-height: 32px;
  padding: 0;
}

.icon {
  width: 18px;
  height: 18px;
  fill: none;
  stroke: currentColor;
  stroke-width: 2.1;
  stroke-linecap: round;
  stroke-linejoin: round;
}


.sr-only {
  position: absolute;
  width: 1px;
  height: 1px;
  padding: 0;
  margin: -1px;
  overflow: hidden;
  clip: rect(0, 0, 0, 0);
  white-space: nowrap;
  border: 0;
}

.operon-table-actions {
  display: flex;
  flex-wrap: wrap;
  align-items: flex-start;
  justify-content: space-between;
  gap: 10px;
}

.table-actions {
  display: flex;
  flex-wrap: wrap;
  align-items: flex-start;
  justify-content: space-between;
  gap: 10px;
}

.search-form {
  display: flex;
  flex-wrap: wrap;
  gap: 8px;
  align-items: center;
  padding-bottom: 12px;
}

.search-form input {
  width: min(420px, calc(100vw - 72px));
  min-height: 34px;
  border: 1px solid var(--border);
  border-radius: 2px;
  background: var(--surface);
  color: var(--text);
  font: inherit;
  font-size: 14px;
  padding: 6px 9px;
}

.filters {
  position: relative;
}

.filters-button {
  display: inline-flex;
  align-items: center;
  gap: 6px;
  min-height: 32px;
  border: 0;
  background: transparent;
  color: var(--muted);
}

.filters-button:hover {
  color: var(--text);
}

.filters-menu {
  position: absolute;
  z-index: 10;
  top: calc(100% + 8px);
  left: 0;
  display: none;
  width: min(420px, calc(100vw - 72px));
  padding: 18px;
  border: 1px solid var(--border);
  border-radius: 2px;
  background: var(--surface);
}

.filters-menu.open {
  display: block;
}

.filters-close {
  position: absolute;
  top: 8px;
  right: 8px;
  width: 30px;
  min-width: 30px;
  min-height: 30px;
  padding: 0;
  border: 0;
  background: transparent;
}

.filters-grid {
  display: grid;
  gap: 12px;
  padding-top: 16px;
}

.filters-grid label {
  display: grid;
  gap: 5px;
  color: var(--muted);
  font-size: 12px;
  font-weight: 700;
  text-transform: uppercase;
}

.filters-grid input,
.filters-grid select {
  width: 100%;
  min-height: 34px;
  border: 1px solid var(--border);
  border-radius: 2px;
  background: var(--surface);
  color: var(--text);
  font: inherit;
  font-size: 14px;
  text-transform: none;
}

.filters-grid input {
  padding: 6px 9px;
}

.filters-grid select {
  padding: 6px 8px;
}

.filters-grid .toggle-filter {
  display: flex;
  align-items: center;
  gap: 8px;
}

.filters-grid .toggle-filter input {
  width: 16px;
  min-height: 16px;
  padding: 0;
}

.filters-actions {
  display: flex;
  justify-content: flex-end;
  gap: 10px;
  margin-top: 16px;
}

.pager {
  display: flex;
  flex-wrap: wrap;
  align-items: center;
  justify-content: flex-end;
  gap: 12px;
  padding: 0 0 12px;
  background: transparent;
}

.pager-button {
  color: var(--accent);
  text-decoration: none;
}

.pager-button:hover {
  color: var(--accent-dark);
}

.pager-current,
.pager-range {
  color: var(--muted);
  font-size: 13px;
  font-weight: 600;
}

.return-link,
.table-link {
  color: var(--accent);
  text-decoration: none;
}

.table-link {
  display: inline-flex;
  align-items: center;
  gap: 3px;
  max-width: 100%;
  vertical-align: baseline;
}

.table-link .link-arrow {
  width: 0.82em;
  height: 0.82em;
  flex: 0 0 auto;
  color: currentColor;
  stroke-width: 2.4;
}

.table-link span {
  min-width: 0;
  overflow-wrap: anywhere;
}

.sort-link {
  display: inline-flex;
  align-items: center;
  gap: 5px;
  color: var(--text);
  text-decoration: none;
}

.sort-link:hover,
.sort-link.active {
  color: var(--accent);
}

.sort-icon {
  width: 14px;
  height: 14px;
}

.return-link {
  display: inline-flex;
  align-items: center;
  gap: 5px;
  color: var(--muted);
  font-size: 14px;
}

.return-link .icon {
  width: 16px;
  height: 16px;
}

.return-link:hover,
.table-link:hover {
  color: var(--accent-dark);
  text-decoration: underline;
}

.gene-viewer {
  overflow-x: auto;
  padding: 0;
}

.gene-viewer-toolbar {
  display: flex;
  flex-wrap: wrap;
  align-items: center;
  justify-content: space-between;
  gap: 12px;
  padding: 12px 14px;
  border-bottom: 1px solid var(--border);
  background: var(--accent-light);
}

.viewer-button {
  min-height: 30px;
  padding: 5px 9px;
  font-size: 13px;
}

.viewer-status {
  color: var(--muted);
  font-size: 13px;
  font-weight: 700;
}

.viewer-status-group {
  display: flex;
  flex-wrap: wrap;
  justify-content: flex-end;
  gap: 10px;
}

.gene-contig-status {
  color: var(--text);
  font-weight: 800;
}

.gene-window {
  padding: 18px 0;
}

.gene-svg {
  display: block;
  width: 100%;
  min-width: 1000px;
  max-width: none;
  height: auto;
}

.genome-viewer {
  overflow-x: auto;
  padding: 0;
}

.genome-window {
  padding: 8px 0 4px;
}

.genome-svg {
  display: block;
  width: 100%;
  min-width: 1000px;
  max-width: none;
  height: auto;
}

.genome-contig-status {
  color: var(--text);
  font-size: 13px;
  font-weight: 800;
}

.genome-occurrence-link {
  cursor: pointer;
}

.genome-occurrence {
  stroke: rgba(255, 255, 255, 0.9);
  stroke-width: 1.2;
}

.genome-occurrence-link:hover .genome-occurrence,
.genome-occurrence-link:focus .genome-occurrence {
  stroke: var(--accent-dark);
  stroke-width: 1.8;
}

.gene-track {
  stroke: #8caabd;
  stroke-width: 1.4;
  stroke-dasharray: 4 5;
}

.gene-arrow {
  stroke: none;
}

.gene-color-0 {
  fill: #2563eb;
}

.gene-color-1 {
  fill: #f97316;
}

.gene-color-2 {
  fill: #16a34a;
}

.gene-color-3 {
  fill: #7c3aed;
}

.gene-color-4 {
  fill: #0891b2;
}

.gene-color-5 {
  fill: #db2777;
}

.gene-color-6 {
  fill: #4f46e5;
}

.gene-color-7 {
  fill: #ca8a04;
}

.gene-color-8 {
  fill: #0d9488;
}

.gene-color-9 {
  fill: #dc2626;
}

.genome-occurrence-color-0 {
  fill: #2563eb;
}

.genome-occurrence-color-1 {
  fill: #f97316;
}

.genome-occurrence-color-2 {
  fill: #16a34a;
}

.genome-occurrence-color-3 {
  fill: #7c3aed;
}

.genome-occurrence-color-4 {
  fill: #0891b2;
}

.genome-occurrence-color-5 {
  fill: #db2777;
}

.genome-occurrence-color-6 {
  fill: #4f46e5;
}

.genome-occurrence-color-7 {
  fill: #ca8a04;
}

.genome-occurrence-color-8 {
  fill: #0d9488;
}

.genome-occurrence-color-9 {
  fill: #dc2626;
}

.gene-label {
  fill: #17212b;
  font-size: 8px;
  font-weight: 400;
  letter-spacing: 0;
  pointer-events: none;
}

.gene-label-reverse {
  fill: #334155;
}

.empty,
.error,
.loading {
  padding: 20px;
  border: 1px solid var(--border);
  border-radius: 2px;
  background: var(--surface);
}

.error {
  border-color: #fecdd3;
  color: var(--danger);
}

@media (max-width: 820px) {
  .sidebar {
    position: static;
    width: 100%;
    padding: 14px;
  }

  .nav {
    grid-template-columns: repeat(4, minmax(0, 1fr));
    margin-top: 14px;
  }

  .nav a {
    text-align: center;
  }

  .main {
    margin-left: 0;
    padding: 20px;
  }

}

@media (max-width: 560px) {
  .main {
    padding: 16px;
  }

  th,
  td {
    padding: 10px;
  }

  .gene-viewer-toolbar {
    align-items: flex-start;
  }

  .gene-viewer-toolbar .button-row,
  .viewer-status-group {
    width: 100%;
  }

  .viewer-status-group {
    justify-content: flex-start;
  }
}